Max Lloyd
Max Lloyd's research interests include isotope geochemistry, biogeochemical cycles, plant metabolism, and surface and near-surface carbon cycling.
In the News
![Research finds increasing temperatures can limit ability of trees to filter air: '[Trees] are essentially coughing instead of breathing'](/sites/default/files/styles/medium_4_3_/public/cover-images/research-finds-increasing-temperatures-can-limit-ability-trees-filter-air-trees-are-essentially.jpg?h=d1cb525d&itok=KqDN2eW5)
Research Keywords
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Recent Publications
A survey of methoxyl and methyl isotopic clumping in a variety of biologic and synthetic materials
Sprengel, R. S., Lloyd, M. K. & Stolper, D. A., Mar 5 2026, In: Chemical Geology. 703, 123243.Research output: Contribution to journal › Article › peer-review
Temperature Sensitivity of Leaf Wax n-Alkane Carbon Isotopes: Mechanisms and Implications for Terrestrial Paleoclimates
Xie, H., Lloyd, M. K., Baczynski, A. A., Diefendorf, A. F. & Freeman, K. H., Feb 16 2026, In: Geophysical Research Letters. 53, 3, e2025GL119656.Research output: Contribution to journal › Letter › peer-review
Modulation of methyl–coenzyme M reductase expression alters the isotopic composition of microbial methane
Gropp, J., Bill, M., Lloyd, M. K., Stein, R. A., Nayak, D. D. & Stolper, D. A., Aug 14 2025, In: Science. 389, 6761, p. 711-715 5 p.Research output: Contribution to journal › Article › peer-review
Agricultural Management and Culinary Culture in the Late Bronze Age and Early Iron Age Aegean: Archaeobotanical and Stable Isotope Insights from Region I, Xeropolis, Lefkandi
Livarda, A., Kotzamani, G., Diffey, C., Evely, D., Sabato, D., Lloyd, M. & Lemos, I., 2025, (Accepted/In press) In: Environmental Archaeology.Research output: Contribution to journal › Article › peer-review
Isotopic evidence for elevated photorespiration during the last glacial period
Lloyd, M. K., Sprengel, R. S., Wortham, B. E., Dunn, R. E., Ibarra, D. E., Dawson, T. E. & Stolper, D. A., Dec 2025, In: Nature Geoscience. 18, 12, p. 1232-1238 7 p.Research output: Contribution to journal › Article › peer-review
Discordance dating: A new approach for dating alteration events
Reimink, J. R., Beckman, R., Schoonover, E., Lloyd, M., Garber, J., Davies, J. H. F. L., Cerminaro, A., Perrot, M. G. & Smye, A., Aug 19 2025, In: Geochronology. 7, 3, p. 369-385 17 p.Research output: Contribution to journal › Article › peer-review
Anaerobic oxidation of methane from abandoned oil and gas wells leaking into aquifers
Shaheen, S. W., Lloyd, M. K., Roden, E. E. & Brantley, S. L., Nov 2025, In: Geochimica et Cosmochimica Acta. 408, p. 269-282 14 p.Research output: Contribution to journal › Article › peer-review
Temperatures of Vein Formation Associated With Plate Interface Deformation Constrained by Oxygen and Clumped Isotope Thermometry
Chen, T. W., Smye, A., Lloyd, M., Fisher, D. & Hashimoto, Y., Jun 2024, In: Geochemistry, Geophysics, Geosystems. 25, 6, e2024GC011516.Research output: Contribution to journal › Article › peer-review
Microbial Autotrophy Recorded by Carbonate Dual Clumped Isotope Disequilibrium
Ingalls, M., Leapaldt, H. C. & Lloyd, M. K., Jun 2024, In: Geochemistry, Geophysics, Geosystems. 25, 6, e2024GC011590.Research output: Contribution to journal › Article › peer-review
Isotopic clumping in wood as a proxy for photorespiration in trees
Lloyd, M. K., Stein, R. A., Ibarra, D. E., Barclay, R. S., Wing, S. L., Stahle, D. W., Dawson, T. E. & Stolper, D. A., 2023, In: Proceedings of the National Academy of Sciences of the United States of America. 120, 46, 2306736120.Research output: Contribution to journal › Article › peer-review
Triple oxygen isotope systematics of diagenetic recrystallization of diatom opal-A to opal-CT to microquartz in deep sea sediments
Ibarra, D. E., Yanchilina, A. G., Lloyd, M. K., Methner, K. A., Chamberlain, C. P., Yam, R., Shemesh, A. & Stolper, D. A., Mar 1 2022, In: Geochimica et Cosmochimica Acta. 320, p. 304-323 20 p.Research output: Contribution to journal › Article › peer-review
Constraints on Early Paleozoic Deep-Ocean Oxygen Concentrations From the Iron Geochemistry of the Bay of Islands Ophiolite
Stolper, D. A., Pu, X., Lloyd, M. K., Christensen, N. I., Bucholz, C. E. & Lange, R. A., Jul 2022, In: Geochemistry, Geophysics, Geosystems. 23, 7, e2021GC010196.Research output: Contribution to journal › Article › peer-review
Clumped 13CH2D and 12CHD2 compositions of methyl groups from wood and synthetic monomers: Methods, experimental and theoretical calibrations, and initial results
Lloyd, M. K., Eldridge, D. L. & Stolper, D. A., Mar 15 2021, In: Geochimica et Cosmochimica Acta. 297, p. 233-275 43 p.Research output: Contribution to journal › Article › peer-review
Methoxyl stable isotopic constraints on the origins and limits of coal-bed methane
Lloyd, M. K., Trembath-Reichert, E., Dawson, K. S., Feakins, S. J., Mastalerz, M., Orphan, V. J., Sessions, A. L. & Eiler, J. M., Nov 12 2021, In: Science. 374, 6569, p. 894-897 4 p.Research output: Contribution to journal › Article › peer-review
Carbonate clumped isotope constraints on burial, uplift and exhumation histories of the Colorado Plateau
Ryb, U., Lloyd, M. K. & Eiler, J. M., Jul 15 2021, In: Earth and Planetary Science Letters. 566, 116964.Research output: Contribution to journal › Article › peer-review