Max Lloyd's research interests include isotope geochemistry, biogeochemical cycles, plant metabolism, and surface and near-surface carbon cycling.
In the News
![Research finds increasing temperatures can limit ability of trees to filter air: '[Trees] are essentially coughing instead of breathing'](/sites/default/files/styles/medium_4_3_/public/cover-images/research-finds-increasing-temperatures-can-limit-ability-trees-filter-air-trees-are-essentially.jpg?h=d1cb525d&itok=KqDN2eW5)
Research Keywords
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Recent Publications
Modulation of methyl–coenzyme M reductase expression alters the isotopic composition of microbial methane
Gropp, J., Bill, M., Lloyd, M. K., Stein, R. A., Nayak, D. D. & Stolper, D. A., Aug 14 2025, In: Science. 389, 6761, p. 711-715 5 p.Research output: Contribution to journal › Article › peer-review
Agricultural Management and Culinary Culture in the Late Bronze Age and Early Iron Age Aegean: Archaeobotanical and Stable Isotope Insights from Region I, Xeropolis, Lefkandi
Livarda, A., Kotzamani, G., Diffey, C., Evely, D., Sabato, D., Lloyd, M. & Lemos, I., 2025, (Accepted/In press) In: Environmental Archaeology.Research output: Contribution to journal › Article › peer-review
Isotopic evidence for elevated photorespiration during the last glacial period
Lloyd, M. K., Sprengel, R. S., Wortham, B. E., Dunn, R. E., Ibarra, D. E., Dawson, T. E. & Stolper, D. A., Dec 2025, In: Nature Geoscience. 18, 12, p. 1232-1238 7 p.Research output: Contribution to journal › Article › peer-review
Discordance dating: A new approach for dating alteration events
Reimink, J. R., Beckman, R., Schoonover, E., Lloyd, M., Garber, J., Davies, J. H. F. L., Cerminaro, A., Perrot, M. G. & Smye, A., Aug 19 2025, In: Geochronology. 7, 3, p. 369-385 17 p.Research output: Contribution to journal › Article › peer-review
Anaerobic oxidation of methane from abandoned oil and gas wells leaking into aquifers
Shaheen, S. W., Lloyd, M. K., Roden, E. E. & Brantley, S. L., Nov 2025, In: Geochimica et Cosmochimica Acta. 408, p. 269-282 14 p.Research output: Contribution to journal › Article › peer-review
Temperatures of Vein Formation Associated With Plate Interface Deformation Constrained by Oxygen and Clumped Isotope Thermometry
Chen, T. W., Smye, A., Lloyd, M., Fisher, D. & Hashimoto, Y., Jun 2024, In: Geochemistry, Geophysics, Geosystems. 25, 6, e2024GC011516.Research output: Contribution to journal › Article › peer-review
Microbial Autotrophy Recorded by Carbonate Dual Clumped Isotope Disequilibrium
Ingalls, M., Leapaldt, H. C. & Lloyd, M. K., Jun 2024, In: Geochemistry, Geophysics, Geosystems. 25, 6, e2024GC011590.Research output: Contribution to journal › Article › peer-review
Isotopic clumping in wood as a proxy for photorespiration in trees
Lloyd, M. K., Stein, R. A., Ibarra, D. E., Barclay, R. S., Wing, S. L., Stahle, D. W., Dawson, T. E. & Stolper, D. A., 2023, In: Proceedings of the National Academy of Sciences of the United States of America. 120, 46, 2306736120.Research output: Contribution to journal › Article › peer-review
Triple oxygen isotope systematics of diagenetic recrystallization of diatom opal-A to opal-CT to microquartz in deep sea sediments
Ibarra, D. E., Yanchilina, A. G., Lloyd, M. K., Methner, K. A., Chamberlain, C. P., Yam, R., Shemesh, A. & Stolper, D. A., Mar 1 2022, In: Geochimica et Cosmochimica Acta. 320, p. 304-323 20 p.Research output: Contribution to journal › Article › peer-review
Constraints on Early Paleozoic Deep-Ocean Oxygen Concentrations From the Iron Geochemistry of the Bay of Islands Ophiolite
Stolper, D. A., Pu, X., Lloyd, M. K., Christensen, N. I., Bucholz, C. E. & Lange, R. A., Jul 2022, In: Geochemistry, Geophysics, Geosystems. 23, 7, e2021GC010196.Research output: Contribution to journal › Article › peer-review
Clumped 13CH2D and 12CHD2 compositions of methyl groups from wood and synthetic monomers: Methods, experimental and theoretical calibrations, and initial results
Lloyd, M. K., Eldridge, D. L. & Stolper, D. A., Mar 15 2021, In: Geochimica et Cosmochimica Acta. 297, p. 233-275 43 p.Research output: Contribution to journal › Article › peer-review
Methoxyl stable isotopic constraints on the origins and limits of coal-bed methane
Lloyd, M. K., Trembath-Reichert, E., Dawson, K. S., Feakins, S. J., Mastalerz, M., Orphan, V. J., Sessions, A. L. & Eiler, J. M., Nov 12 2021, In: Science. 374, 6569, p. 894-897 4 p.Research output: Contribution to journal › Article › peer-review
Carbonate clumped isotope constraints on burial, uplift and exhumation histories of the Colorado Plateau
Ryb, U., Lloyd, M. K. & Eiler, J. M., Jul 15 2021, In: Earth and Planetary Science Letters. 566, 116964.Research output: Contribution to journal › Article › peer-review
Triple oxygen isotopes of meteoric hydrothermal systems - implications for palaeoaltimetry
Chamberlain, C. P., Ibarra, D. E., Lloyd, M. K., Kukla, T., Sjostrom, D., Gao, Y. & Sharp, Z. D., 2020, In: Geochemical Perspectives Letters. 15, p. 6-9 4 p.Research output: Contribution to journal › Article › peer-review
Massive formation of early diagenetic dolomite in the Ediacaran ocean: Constraints on the "dolomite problem"
Chang, B., Chang, B., Li, C., Liu, D., Foster, I., Tripati, A., Tripati, A., Tripati, A., Lloyd, M. K., Maradiaga, I., Maradiaga, I., Luo, G., An, Z., She, Z., Xie, S., Tong, J., Huang, J., Algeo, T. J., Algeo, T. J. & Algeo, T. J. & 2 others, , Jun 23 2020, In: Proceedings of the National Academy of Sciences of the United States of America. 117, 25, p. 14005-14014 10 p.Research output: Contribution to journal › Article › peer-review