Timothy Ryan

IEE Faculty Member
Titles and Affiliations
Professor, Anthropology

Research Summary

Functional morphology, bone biomechanics, primate locomotion, primate and human evolution, high-resolution computed tomography, three-dimensional scientific visualization.

In the News

Research Keywords

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Recent Publications

Becoming adults: exploring the late ontogeny of the human talus

Figus, C., Sorrentino, R., Carlson, K. J., Colombo, A., Bortolini, E., Bernardini, F., Erjavec, I., Hajdu, T., Szeniczey, T., Mateovics-László, O., Pap, I., Tuniz, C., Novak, M., Belcastro, M. G., Ryan, T. M. & Benazzi, S., 2023, In: Frontiers in Ecology and Evolution. 11, 1205277.

Research output: Contribution to journalArticlepeer-review

Morphologies in-between: The impact of the first steps on the human talus

Figus, C., Stephens, N. B., Sorrentino, R., Bortolini, E., Arrighi, S., Higgins, O. A., Lugli, F., Marciani, G., Oxilia, G., Romandini, M., Silvestrini, S., Baruffaldi, F., Belcastro, M. G., Bernardini, F., Festa, A., Hajdu, T., Mateovics-László, O., Pap, I., Szeniczey, T., Tuniz, C., & 2 othersRyan, T. M. & Benazzi, S., Jan 2023, In: Anatomical Record. 306, 1, p. 124-142 19 p.

Research output: Contribution to journalArticlepeer-review

Morphological and evolutionary insights into the keystone element of the human foot’s medial longitudinal arch

Sorrentino, R., Carlson, K. J., Orr, C. M., Pietrobelli, A., Figus, C., Li, S., Conconi, M., Sancisi, N., Belvedere, C., Zhu, M., Fiorenza, L., Hublin, J. J., Jashashvili, T., Novak, M., Patel, B. A., Prang, T. C., Williams, S. A., Saers, J. P. P., Stock, J. T., Ryan, T., & 6 othersMyerson, M., Leardini, A., DeSilva, J., Marchi, D., Belcastro, M. G. & Benazzi, S., Dec 2023, In: Communications Biology. 6, 1, 1061.

Research output: Contribution to journalArticlepeer-review

Human talar ontogeny: Insights from morphological and trabecular changes during postnatal growth

Figus, C., Stephens, N. B., Sorrentino, R., Bortolini, E., Arrighi, S., Lugli, F., Marciani, G., Oxilia, G., Romandini, M., Silvestrini, S., Baruffaldi, F., Belcastro, M. G., Bernardini, F., Erjavec, I., Festa, A., Hajdu, T., Mateovics-László, O., Novak, M., Pap, I., Szeniczey, T., & 3 othersTuniz, C., Ryan, T. M. & Benazzi, S., Oct 2022, In: American Journal of Biological Anthropology. 179, 2, p. 211-228 18 p.

Research output: Contribution to journalArticlepeer-review

Ontogenetic Patterning of Human Subchondral Bone Microarchitecture in the Proximal Tibia

Goliath, J. R., Gosman, J. H., Stout, S. D. & Ryan, T. M., Jul 2022, In: Biology. 11, 7, 1002.

Research output: Contribution to journalArticlepeer-review

South-to-north migration preceded the advent of intensive farming in the Maya region

Kennett, D. J., Lipson, M., Prufer, K. M., Mora-Marín, D., George, R. J., Rohland, N., Robinson, M., Trask, W. R., Edgar, H. H. J., Hill, E. C., Ray, E. E., Lynch, P., Moes, E., O’Donnell, L., Harper, T. K., Kate, E. J., Ramos, J., Morris, J., Gutierrez, S. M., Ryan, T. M., & 3 othersCulleton, B. J., Awe, J. J. & Reich, D., Dec 2022, In: Nature communications. 13, 1, 1530.

Research output: Contribution to journalArticlepeer-review

Growth and development of trabecular structure in the calcaneus of Japanese macaques (Macaca fuscata) reflects locomotor behavior, life history, and neuromuscular development

Saers, J. P. P., Gordon, A. D., Ryan, T. M. & Stock, J. T., Jul 2022, In: Journal of Anatomy. 241, 1, p. 67-81 15 p.

Research output: Contribution to journalArticlepeer-review

Trabecular bone ontogeny tracks neural development and life history among humans and non-human primates

Saers, J. P. P., Gordon, A. D., Ryan, T. M. & Stock, J. T., Dec 6 2022, In: Proceedings of the National Academy of Sciences of the United States of America. 119, 49, e2208772119.

Research output: Contribution to journalArticlepeer-review

The talar morphology of a hypochondroplasic dwarf: A case study from the Italian Late Antique period

Sorrentino, R., Carlson, K. J., Figus, C., Pietrobelli, A., Stephens, N. B., DeMars, L. J. D., Saers, J. P. P., Armando, J., Bettuzzi, M., Guarnieri, T., Oxilia, G., Vazzana, A., Parr, W., Turley, K., Morigi, M. P., Stock, J. T., Ryan, T. M., Benazzi, S., Marchi, D. & Belcastro, M. G., Mar 1 2022, In: International Journal of Osteoarchaeology. 32, 2, p. 429-443 15 p.

Research output: Contribution to journalArticlepeer-review

Potential evolutionary body size reduction in a Malagasy primate (Propithecus verreauxi) in response to human size-selective hunting pressure

Sullivan, A. P., Godfrey, L. R., Lawler, R. R., Randrianatoandro, H., Eccles, L., Culleton, B., Ryan, T. M. & Perry, G. H., Jul 2022, In: American Journal of Biological Anthropology. 178, 3, p. 385-398 14 p.

Research output: Contribution to journalArticlepeer-review

Using point clouds to investigate the relationship between trabecular bone phenotype and behavior: An example utilizing the human calcaneus

DeMars, L. J. D., Stephens, N. B., Saers, J. P. P., Gordon, A., Stock, J. T. & Ryan, T. M., Mar 1 2021, In: American Journal of Human Biology. 33, 2, e23468.

Research output: Contribution to journalArticlepeer-review

New Pliocene hominin remains from the Leado Dido'a area of Woranso-Mille, Ethiopia

Melillo, S. M., Gibert, L., Saylor, B. Z., Deino, A., Alene, M., Ryan, T. M. & Haile-Selassie, Y., Apr 2021, In: Journal of Human Evolution. 153, 102956.

Research output: Contribution to journalArticlepeer-review

Automated resolution independent method for comparing in vivo and dry trabecular bone

Saers, J. P. P., DeMars, L. J., Stephens, N. B., Jashashvili, T., Carlson, K. J., Gordon, A. D., Ryan, T. M. & Stock, J. T., Apr 2021, In: American Journal of Physical Anthropology. 174, 4, p. 822-831 10 p.

Research output: Contribution to journalArticlepeer-review

Combinations of trabecular and cortical bone properties distinguish various loading modalities between athletes and controls

Saers, J. P. P., DeMars, L. J., Stephens, N. B., Jashashvili, T., Carlson, K. J., Gordon, A. D., Shaw, C. N., Ryan, T. M. & Stock, J. T., Mar 2021, In: American Journal of Physical Anthropology. 174, 3, p. 434-450 17 p.

Research output: Contribution to journalArticlepeer-review

Unique foot posture in Neanderthals reflects their body mass and high mechanical stress

Sorrentino, R., Stephens, N. B., Marchi, D., DeMars, L. J. D., Figus, C., Bortolini, E., Badino, F., Saers, J. P. P., Bettuzzi, M., Boschin, F., Capecchi, G., Feletti, F., Guarnieri, T., May, H., Morigi, M. P., Parr, W., Ricci, S., Ronchitelli, A., Stock, J. T., Carlson, K. J., & 3 othersRyan, T. M., Belcastro, M. G. & Benazzi, S., Dec 2021, In: Journal of Human Evolution. 161, 103093.

Research output: Contribution to journalArticlepeer-review